The evolutionary history of Fagus in western Eurasia: Evidence from genes, morphology and the fossil record

By Thomas Denk, G. Grimm, K. Stögerer, M. Langer, V. Hemleben. Plant Systematics and Evolution, 2002

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Biological Flora of the British Isles: Fagus sylvatica (2012, background): “More recently, Denk et al. (2002), after investigating nuclear rDNA internal transcribed spacer sequences, concluded that there is only a single species, Fagus sylvatica, in Europe and Asia Minor.”
Palaeoclimatic models help to understand current distribution of Caucasian forest species (2012, methodology): “caucasica, respectively (Farjon, 1990; Veith et al., 1998; Denk et al., 2002; Schütt, 2005), and were used as their ecological counterparts in Europe.”
Phylogeny of Fagus L. (Fagaceae) based on morphological data (2003, methodology): “The F. engleriana group of Zetter (1984) corresponds to the subgenus Engleriana Shen that is also recognised in the present study and by Denk et al. (2002).”