Plant species diversity and environmental heterogeneity: spatial scale and competing hypotheses

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Scale- and taxon-dependent biodiversity patterns of dry grassland vegetation in Transylvania (2014, background): “The heterogeneity-diversity hypothesis (HDR) assumes that heterogeneous environments can host more species because of more available niches (see Lundholm, 2009; Tamme et al., 2010).”
Linking small-scale topography with microclimate, plant species diversity and intra-specific trait variation in an alpine landscape (2015, background): “The relationship between environmental heterogeneity and plant species diversity has been extensively studied, and there is now abundant evidence suggesting that an increase in habitat heterogeneity increases the number of co-existing species (reviewed in Lundholm 2009).”