Sexual selection and horn allometry in the dung beetle Euoniticellus intermedius

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Horn size predicts physical performance in the beetle Euoniticellus intermedius (Coleoptera: Scarabaeidae) (2005, background): “All insects were taken from the same E. intermedius laboratory culture used in Pomfret & Knell (2005) , originally started with >100 adult beetles collected from South Africa.”
On the analysis of non‐linear allometries (2009, background): “However other mechanisms may also contribute to this, such as constraints on horn length or an uncoupling of horn length and body size ( Pomfret & Knell, 2006 ).”
A functional perspective on sexual selection: insights and future prospects (2006, background): “Victory in fights between male horned dung beetles, Euoniticellus intermedius, is determined by size-adjusted horn size (Pomfret & Knell 2006), which is also correlated with two types of performance abilities used by males during fights (Lailvaux et al. 2005).”
The Evolution of Animal Weapons (2008, background): “Both body size and horn length confer significant performance advantages to fighting males (76, 185, 198, 204, 215), and this can translate into fertilization success (124).”