Aluminium tolerance is achieved by exudation of citric acid from roots of soybean (Glycine max)

By Zhen-ming Yang, M. Sivaguru, W. Horst, H. Matsumoto. 2000

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Aluminum hyperaccumulation in angiosperms: A review of its phylogenetic significance (2002, background): “…of aluminum-chelating ligands (organic acids) from the root apex, such as citrate in Cassia tora and soybean (Glycine max, Ma et al., 1997b; Yang et al., 2000), oxalate in buckwheat (Fagopyrum esculentum, Ma et al., 1997c; Zheng et al., 1998), and citrate and malate in Arabidopsis mutants…”
Physiology and molecular biology of stress tolerance in plants (2006, background): “Some plant species achieve Al tolerance by Al-activated exudation of organic acids from roots (Yang et al., 2000; Ryan et al., 2001).”
Effect of Al in Soil on Photosynthesis and Related Morphological and Physiological Characteristics of Two Soybean Genotypes (2007, background): “In addition, Yang et al. (2000, 2003) reported that photosynthesis plays an important role in the exudation of citrate exudation, which is an important Al tolerance mechanism in aluminum-tolerant soybean plant, but the contribution of shoots combined with light to Al-induced secretion of citrate…”