Why do male Callosobruchus maculatus harm their mates

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Female Callosobruchus maculatus mate when they are thirsty: resource-rich ejaculates as mating effort in a beetle (2007, background): “Previous studies of C. maculatus have also demonstrated costs of mating caused by the spines on male genitalia (Crudgington & Siva-Jothy 2000; Edvardsson & Tregenza 2005 ) and have shown that females reduce their remating propensity when they receive relatively large ejaculates (Eady 1995; Savalli ”
Do males matter? The role of males in population dynamics (2007, background): “…found that the spines may have evolved as a side-adaptation, rather than to aid males in sperm competition, demonstrated by the fact that preventing females from kicking during their second copulation did not make the second mates more successful in sperm competition (Edvardsson and Tregenza 2005).”
The effects of copulation duration in the bruchid beetle Callosobruchus maculatus (2006, methodology): “maculatus transfer large ejaculates that, in virgin males, can weigh up to 10% of their body weight (Edvardsson and Tregenza, 2005; Savalli and Fox, 1998).”