Interspecific competition in tree squirrels: do introduced grey squirrels (Sciurus carolinensis) deplete tree seeds hoarded by red squirrels (S. vulgaris)?

By L. Wauters, Guido Tosi, J. Gurnell. Behavioral Ecology and Sociobiology, 2002

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Alien species and interspecific competition: effects of introduced eastern grey squirrels on red squirrel population dynamics (2004, background): “We showed that in coexisting populations in northern Italy, grey squirrels pilfering red squirrels’ food caches, caused a reduced energy intake in red squirrels with a high degree of interspecific core-area overlap, and a decrease of body mass in spring (Wauters et al. 2002a).”
A grey future for Europe: Sciurus carolinensis is replacing native red squirrels in Italy (2013, background): “This replacement is caused by competition for food resources between the two species that reduces juvenile body growth and recruitment, and female breeding success in red squirrel (Wauters et al. 2002a, b; Gurnell et al. 2004).”
Interspecific competition between native Eurasian red squirrels and alien grey squirrels: does resource partitioning occur? (2002, background): “Red squirrels fed intensively on walnuts and black walnuts in autumn/winter and cached large numbers of them for use in late winter/spring ( Wauters et al. 2001, 2002 ).”
Competitive exclusion after invasion? (2008, background): “in habitat or diet) (Nilsson 1967, 1978) (ii) density reduction or extinction (negative population growth) (Hardin 1960) (iii) reduced individual growth (Diehl and Eklov 1995) (iv) reduced food intake (Bohn and Amundsen 2001; Wauters et al. 2002 ) (v) altered community composition of prey towards s”