Oxidative Stress in the Frozen Plant: A Free Radical Point of View

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Unifying perspectives of some mechanisms basic to desiccation tolerance across life forms (2006, background): “While superoxide itself is relatively short lived, it may dismutate to form H2O2 which, if not reduced to water, can be involved in a major route of †OH formation by the Fenton reaction, Fe2þ þ H2O2 ! Fe3þ þ †OH þ HO(2) (Hendry, 1993; Benson and Bremner, 2004).”
Physiology of Desiccation-Sensitive (Recalcitrant) Seeds and the Implications for Cryopreservation (2013, background): “ROS are normal components of aerobic metabolism, but their production is enhanced under conditions of stress, inter alia wounding, desiccation, and chilling/freezing stress (reviewed by Smirnoff 1993; Walters et al. 2002; Benson and Bremner 2004).”
Cryopreservation of Phytodiversity: A Critical Appraisal of Theory & Practice (2008, background): “The readers are directed to Benson (2008), Benson and Bremner (2004), Mazur (2004), Muldrew et al. (2004) and Taylor et al. (2004) for reviews on more fundamental aspects.”
Current Issues in Plant Cryopreservation (2012, background): “The electron transport chain used in respiration and photosynthesis are the major producers of ROS, caused by the leakage of free electrons onto molecular oxygen (O2), resulting in the formation of superoxide (Benson & Bremner, 2004; Benson, 1990; Halliwell & Gutteridge, 2007).”