Relationships between Large―Spike Phenotype, Grain Number, and Yield Potential in Spring Wheat

By O. Gaju, Matthew P Reynolds, D. Sparkes, M. J. Foulkes. 2009

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Effects of Drought Stress on Some Agronomic and Morpho-Physiological Traits in Durum Wheat Genotypes (2020, background): “Indeed, grain yield can be increased through manipulation of yield-related traits such as spike length, number of spikelets per spike, and grains per spike [20].”
The Physiological Basis of the Genetic Progress in Yield Potential of CIMMYT Spring Wheat Cultivars from 1966 to 2009 (2015, result): “…grains per square meter and grain weight has been frequently observed among wheat genotypes (Foulkes et al., 2011) and a negative relationship between SPI and FE in wheat has also previously been reported in some studies (e.g., Gaju et al., 2009; González et al., 2011; Lázaro and Abbate, 2012).”
Raising yield potential of wheat. III. Optimizing partitioning to grain while maintaining lodging resistance. (2011, background): “There is generally a negative relationship between these two determinants of grain number per unit area (Gaju et al., 2009; Foulkes et al., 2009b) and it will be crucial to identify opportunities to break this negative relationship.”