Complementary biocontrol of aphids by the ladybird beetle Harmonia axyridis and the parasitoid Aphelinus asychis on greenhouse roses

By W. E. Snyder, Stacey Ballard, Suanne Yang, Garrett M. Clevenger, T. Miller, J. Ahn, Timothy D. Hatten, A. Berryman. 2004

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Impact of intraguild predation by adult Harmonia axyridis (Coleoptera: Coccinellidae) on Aphis glycines (Hemiptera: Aphididae) biological control in cage studies (2007, unknown): “…2001; Snyder et al., 2004a,b; Colunga-Garcia and Gage, 1998; Phoofolo and Obrycki, 1998; Cottrell, 2005), inclusion of these predators often results in increased biological control (Colfer and Rosenheim, 2001; Cardinale et al., 2003; Snyder et al., 2004a; Aquilino et al., 2005; Costamagna, 2006).”
Ecology and biological control application of multicoloured Asian ladybird, Harmonia axyridis: A review (2006, methodology): “As an intraguild predator, H. axyridis captured and killed larvae of two native species over 70% of the time, while as a prey, it was captured and killed by the superior predator only 25% of the time (Snyder et al. 2004a).”