Evolution of the mane and group-living in the lion ( Panthera leo ): a review

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Extinction chronology of the cave lion Panthera spelaea (2011, unknown): “However, this interpretation is disputed by Yamaguchi et al. (2004).”
Isotopic evidence for dietary ecology of cave lion (Panthera spelaea) in North-Western Europe: Prey choice, competition and implications for extinction (2011, background): “…cave lions as documented by prehistoric parietal art that would indicate solitary hunting behaviour or hunting by breeding pairs of lion and lioness (Guthrie,1990), although the relationship between mane development and social behaviour of lion is contested by others (e.g., Yamaguchi et al., 2004).”
Did Neanderthals and other early humans sing? Seeking the biological roots of music in the territorial advertisements of primates, lions, hyenas, and wolves (2009, background): “Based on analyses of lion paleoecology, sociobiology, and depictions in cave art, Yamaguchi et al. (2004) conclude Pleistocene lions were group-living, so their behaviour should resemble modern lions.”