Development and distribution of predators and parasitoids during two consecutive years of an Ips typographus (Col., Scolytidae) infestation

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Research on Parasitoids and Predators of Scolytidae – A Review (2007, methodology): “To better evaluate the impact of parasitoids and predators on bark beetle populations, various methods have been used, such as consumption rate based assessments (e.g. Dippel et al. 1997; Wermelinger 2002), life table analyses, and natural enemy exclusion experiments.”
Simulation and analysis of outbreaks of bark beetle infestations and their management at the stand level (2011, background): “respectively, of broken and uprooted trees (Wermelinger, 2002) and thus led to the favoured conditions for the ensuing calamities caused by I.”
Ecology and management of the spruce bark beetle Ips typographus—a review of recent research (2004, background): “In a biennial study, mortality imposed by predators and parasitoids increased from 558 % to 828 % during two years of an ongoing infestation (Wermelinger, 2002).”
Population dynamics of the spruce bark beetle: a long-term study (2013, background): “typographus , a relatively large number of predators and parasitoids are expected to respond numerically to bark beetle density producing negative density feedback on host/prey population (Wermelinger 2002).”