Predator–prey interactions in river networks: comparing shrimp spatial refugia in two drainage basins

By Alan P. Covich, T. Crowl, C. Hein, M. J. Townsend, William H. McDowell. 2009

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Amphidromy in shrimps: a life cycle between rivers and the sea (2013, background): “A major benefit for freshwater shrimps in the headwaters of streams on mountainous tropical islands is that there are few or no fish predators there (e.g., Covich et al., 2009; Blob et al., 2010; Hein et al., 2011).”
Winning the biodiversity arms race among freshwater gastropods: competition and coexistence through shell variability and predator avoidance (2010, background): “They can re-colonize rivers after disturbances such as floods and droughts from these coastal sources and are relatively resilient, dominant consumers in many river food webs (Covich et al., 2006, 2009).”
Biological Diversity in Headwater Streams (2019, background): “The use of headwaters by Atya lanipes and Xiphocaris elongata shrimps in Puerto Rico is interpreted as providing refuge from predation by fishes in downstream reaches [15].”