Effects of gut passage on seed germination: do experiments answer the questions they ask?

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Seed dispersal by Galápagos tortoises (2012, methodology): “Following the recommendations of Samuels & Levey (2005), our germination trials were designed to test whether digesta retention time, the presence of dung in the substrate and the presence of fruit pulp influenced germination patterns of the five focal species.”
Assessing the benefits of frugivory for seed germination: the importance of the deinhibition effect (2006, methodology): “Our data show that the methodological concerns raised in recent papers (Kelly et al. 2004; Samuels & Levey 2005) are valid.”
The dispersal syndrome hypothesis: How animals shaped fruit traits, and how they did not (2020, background): “…such as the number of seeds dispersed (Estrada & Coates-Estrada, 1984; Valenta & Fedigan, 2008), effects of mouth and gut treatment on germination (Samuels & Levey, 2005; Traveset, 1998; Valenta & Fedigan, 2009), seed dispersal distances from parent trees (Hyatt et al., 2003; Levey, Tewksbury, &…”