Weight loss in drone pupae (Apis mellifera) multiply infested by Varroa destructor mites

By Pedro Duay, D. Jong, W. Engels. 2003

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Biology and control of Varroa destructor. (2010, background): “This has also been proven for parasitized drones, which loose 11–19% of their body weight depending of infestation rate (Duay et al., 2003), which led to decreased flight performance (Duay et al., 2002).”
A historical review of managed honey bee populations in Europe and the United States and the factors that may affect them. (2010, background): “Both the adult female and her offspring feed on pupae, where they can cause damage by ingestion of hemolymph, resulting in severe nutritional deficits for the developing bee (Duay et al., 2003; Garedew et al., 2004).”