Length‐specific growth rates in freshwater mussels (Bivalvia: Unionidae): extreme longevity or generalized growth cessation?

By J. Anthony, D. Kesler, W. L. Downing, J. Downing. 2001

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Changing Perspectives on Pearly Mussels, North America's Most Imperiled Animals (2004, background): “However, recent studies of growth rates based on direct measurements of marked individuals in the field suggest that growth rings are not annual and that earlier estimates based on growth rings may underestimate longevity by a factor of 3 to 10 (Anthony et al. 2001).”
Effects of zebra mussels (Dreissena polymorpha) on native bivalves: the beginning of the end or the end of the beginning? (2007, background): “Rings in unionid shells are commonly interpreted as winter growth stops and are used for studies of unionid growth (e.g., Neves and Moyer 1988, Haag and Staton 2003), although some studies have shown that these rings are not always annual (Kesler and Downing 1997, Anthony et al. 2001).”