Relation between oak tree phenology and the secretion of organic matter degrading enzymes by Lactarius quietus ectomycorrhizas before and during bud break

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Decomposers in disguise: mycorrhizal fungi as regulators of soil C dynamics in ecosystems under global change (2008, background): “Courty, Breda & Garbaye (2007) also found that ectomycorrhizal root tips in an old-growth oak forest produce a suite of extracellular enzymes in the early spring that show peak activity immediately before and following bud break.”
Independent roles of ectomycorrhizal and saprotrophic communities in soil organic matter decomposition (2013, background): “Enzyme production by ECM root tips can increase with decreasing access to plant photosynthate (Courty et al., 2007; Courty et al., 2010), suggesting that ECM fungi maymine soil for C when easily-accessible C supplies are low.”
Ectomycorrhizal fungi: the symbiotic route to the root for phosphorus in forest soils (2011, background): “2007), toxic metal pollution (Kieliszewska-Rokicka 1999), rainfall (Courty et al. 2007), nitrogen availability (Kieliszewska-Rokicka 1992; Taniguchi et al.”