Tree species mixing can increase maximum stand density

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Effects of crown architecture and stand structure on light absorption in mixed and monospecific Fagus sylvatica and Pinus sylvestris forests along a productivity and climate gradient through Europe (2018, background): “…that are more productive are likely to have greater stand densities and lower mortality rates; as indicated by higher intercepts of self- thinning lines (Binkley, 1984; Binkley, Senock, Bird, & Cole, 2003; Pretzsch & Biber, 2016; Pretzsch et al., 2015; Reyes- Hernandez, Comeau, & Bokalo, 2013).”
Toward managing mixed-species stands: from parametrization to prescription (2017, background): “2), the vertical layering more heterogeneous (Dănescu et al. 2016, Peck et al. 2014), and the asymmetry of competition higher (Pretzsch et al., 2016a, b), even though growth dominance is similar to monocultures (Pretzsch et al. 2017, Binkley et al. 2006).”
Species interactions increase the temporal stability of community productivity in Pinus sylvestris–Fagus sylvatica mixtures across Europe (2017, unknown): “…on mortality, although significant effects of mixing on tree mortality, self-thinning lines and stand density indices have been reported previously (Binkley 1984, 2003; Condés & Río 2015; Pretzsch & Biber 2016; Woodall, Milles & Vissage 2005), and may influence overyielding as well as stability.”