A model of grid cell development through spatial exploration and spike time-dependent plasticity.

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Layer-Specific Organization of Local Excitatory and Inhibitory Synaptic Connectivity in the Rat Presubiculum (2017, methodology): “Learning rules implementing activitydependent plasticity have already been used to adjust synaptic weights for recurrent synapses between pyramidal cells and interneurons in a model for grid cell development (Widloski and Fiete 2014).”
Attractor and integrator networks in the brain (2021, background): “Given the preserved internal structure and autonomous dynamics of grid cells across states, time, and environments, it follows that various deviations in the spatial tuning curves of grid cells from equilateral grid-like responses in 2- and 3-dimensional spaces [171, 172, 173, 174, 175, 176] likely ”